Ecological Archives A019-008-A1

Tara Hudiburg, Beverly Law, David P. Turner, John L. Campbell, Dan Donato, and Maureen Duane. 2009. Carbon dynamics of Oregon and northern California forests and potential land-based carbon storage. Ecological Applications 19:163–180.

Appendix A. Ecoregion, ownership, and forest type summary tables and statistics.

TABLE A1. Total estimates of biomass, NPP, mortality.

A. Ecoregion total estimates

Ecoregion

Live Biomass Pg C

Dead Biomass Pg C

NPP Pg C yr-1

Mortality Pg C yr-1

BM

0.182

0.056

0.0080

0.0025

CR

0.627

0.119

0.0256

0.0021

EC

0.200

0.042

0.0010

0.0023

KM

0.626

0.088

0.0246

0.0042

SN

0.347

0.066

0.0140

0.0042

WC

0.511

0.116

0.0144

0.0035

CB

0.008

0.001

0.0004

7.6x10-5

CO

0.128

0.140

0.0084

0.0016

CP

0.002

8.7 × 10-5

9.6 × 10-5

1.6 × 10-5

NB

0.006

0.003

0.0003

0.0001

WV

0.073

0.010

0.0034

0.0003

Total

2.710

0.514

0.1091

0.0208

B. Owner total estimates (within ecoregion)

Ecoregion

Live Biomass Pg C

Dead Biomass Pg C

NPP Pg C yr-1

Mortality Pg C yr-1

 

Private

Public

Private

Public

Private

Public

Private

Public

BM

0.035

0.126

0.014

0.038

0.0022

0.0054

0.0004

0.0017

CR

0.284

0.236

0.068

0.040

0.0181

0.0081

0.0016

0.0006

EC

0.059

0.134

0.015

0.028

0.0034

0.0066

0.0006

0.0016

KM

0.163

0.414

0.028

0.057

0.0100

0.0156

0.0010

0.0028

SN

0.088

0.236

0.019

0.044

0.0050

0.0094

0.0011

0.0028

WC

0.070

0.392

0.026

0.087

0.0041

0.0108

0.0004

0.0027

CB

0.000

0.008

-

0.001

1.7 × 10-5

0.0003

4.6 × 10-6

7.2 × 10-5

CO

0.119

0.017

0.014

0.002

0.0081

0.0010

0.0016

0.0002

CP

0.001

0.000

7.3 × 10-5

-

8.1 × 10-5

-

1.4 × 10-5

-

NB

0.002

0.004

-

0.002

9.6 × 10-5

0.0002

1.1 × 10-5

8.4 × 10-5

WV

0.058

0.009

0.007

0.001

0.0031

0.0003

0.0003

2.9 × 10-5

Total

0.880

1.580

0.177

0.300

0.0516

0.0578

0.0070

0.0196

C. Forest types total estimates (within ecoregion)

Ecoregion

Live Biomass Pg C

 

1

2

3

4

5

6

7

Total

BM

0.091

0.008

0.003

0.007

0.067

5.8 × 10-5

0.003

0.179

CR

0.429

0.052

-

0.014

0.002

0.125

0.001

0.623

EC

0.080

5.0 × 10-4

0.003

3.0 × 10-4

0.111

0.004

0.003

0.201

KM

0.478

0.005

-

2.0 × 10-4

0.016

0.126

0.001

0.627

SN

0.256

-

0.003

-

0.055

0.033

0.002

0.348

WC

0.480

0.006

-

0.002

0.016

0.004

0.001

0.508

CB

0.001

-

-

-

0.007

-

4.0 × 10-4

0.008

CO

0.024

0.004

-

-

0.011

0.002

-

0.039

CP

4.0 × 10-4

-

1.0 × 10-4

-

0.001

-

-

0.001

NB

0.002

-

0.001

-

0.001

-

-

0.004

WV

0.060

-

-

-

-

0.012

0.001

0.072

Ecoregion

Dead Biomass Pg C

 

1

2

3

4

5

6

7

Total

BM

0.027

0.004

0.001

0.002

0.015

0.000

0.004

0.052

CR

0.078

0.012

-

0.002

0.001

0.025

4.0 × 10-4

0.117

EC

0.016

1.0 × 10-4

4.0 × 10-4

7.3 × 10-5

0.022

0.001

0.002

0.041

KM

0.067

0.001

-

3.7 × 10-5

0.003

0.014

.001

0.086

SN

0.046

-

4.0 × 10-5

-

0.010

0.004

0.001

0.062

WC

0.107

0.002

-

0.001

0.003

0.001

0.002

0.115

CB

2.0 × 10-5

-

-

-

0.001

-

7.6 × 10-5

0.001

CO

0.003

1.0 × 10-5

-

-

0.002

0.005

-

0.009

CP

=

-

-

-

-

2.2 × 10-5

-

2.2 × 10-5

NB

0.001

-

8.4 × 10-5

-

2.0 × 10-4

-

-

0.001

WV

0.008

-

-

-

-

0.001

3.0 × 10-4

0.009

Ecoregion

NPP Pg C yr-1

 

1

2

3

4

5

6

7

Total

BM

0.0034

0.0003

0.0002

0.0002

0.0036

4.2 × 10-6

0.0002

0.0080

CR

0.0153

0.0024

-

0.0004

0.0001

0.0074

4.3 × 10-5

0.0257

EC

0.0035

2.5 × 10-5

0.0001

4.3 × 10-6

0.0060

0.0002

0.0002

0.0100

KM

0.0165

0.0002

-

6.9 × 10-6

0.0009

0.0069

9.7 × 10-5

0.0247

SN

0.0095

-

0.0001

-

0.0027

0.0016

0.0002

0.0141

WC

0.0131

0.0001

-

5.3 × 10-5

0.0008

0.0002

6.6 × 10-5

0.0144

CB

3.3 × 10-5

-

-

-

0.0003

-

3.7 × 10-5

0.0003

CO

0.0013

0.0001

-

-

0.0008

0.0037

-

0.0059

CP

3.0 × 10-5

-

1.0 × 10-5

-

3.2 × 10-5

-

-

7.2 × 10-5

NB

7.0 × 10-5

-

8.5 × 10-5

-

7.2 × 10

-

1.1 × 10-5

0.0002

WV

0.0026-

-

-

-

-

0.0007

7.6 × 10-5

0.0034

Ecoregion

Mortality Pg C yr-1

 

1

2

3

4

5

6

7

Total

BM

0.0015

0.0001

2.9 × 10-5

9.9 × 10-5

0.0006

1.2 × 10-6

3.9 × 10-5

0.0025

CR

0.0012

0.0002

-

3.3 × 10-5

1.3 × 10-5

0.0007

1.7 × 10-5

0.0021

EC

0.0013

3.6 × 10-6

2.0 × 10-5

2.7 × 10-6

0.0010

3.0 × 10-5

2.4 × 10-5

0.0023

KM

0.0033

3.3 × 10-5

-

1.6 × 10-6

0.0001

0.0007

6.8 × 10-6

0.0042

SN

0.0033

-

2.5 × 10-5

-

0.0005

0.0003

2.5 × 10-5

0.0042

WC

0.0032

3.8 × 10-5

-

2.0 × 10-5

0.0002

2.4 × 10-5

6.5 × 10-6

0.0035

CB

1.0 × 10-5

-

-

-

5.9 × 10-5

-

3.2 × 10-6

0.0001

CO

0.0002

4.2 × 10-6

-

-

7.5 × 10-5

0.0008

-

0.0011

CP

3.7 × 10-6

-

1.1 × 10-6

-

6.0 × 10-6

5.6 × 10-6

-

1.6 × 10-5

NB

6.2 × 10-5

-

1.3 × 10-5

-

1.2 × 10-5

6.7 × 10-7

6.0 × 10-7

8.8 × 10-5

WV

0.0002

-

-

-

-

7.0 × 10-4

3.9 × 10-6

0.0003

1. Forest Types: 1 =Fir/Douglas fir/Hemlock, 2=Cedar/Larch/Sequoia/Redwood, 3=Juniper/Cypress, 4=Spruce, 5=Pine, 6=Hardwoods, 7=Non-Stocked

2. Totals were calculated using the mean value per category and multiplied by the corresponding land area. Note that total regional values will vary by method (i.e., the total sum of each ecoregion estimate vs. the total sum of each ownership within ecoregion estimate will be different because different mean values are used to multiply by disproportionate land areas).


TABLE A2. Ecoregion, owner, and forest type mean estimates (± standard errors) of biomass, NPP, and mortality.

A. Ecoregion mean estimates

Ecoregion

Live Biomass kg C m-2

Dead Biomass kg C m-2

NPP kg C m-2 yr-1

Mortality kg C m-2 yr-1

BM

6.4±0.1

2.0±0.1

0.30±0.00

0.09±0.002

CR

19.1±0.5

3.6±0.1

0.78±0.01

0.05±0.001

EC

6.7±0.1

1.4±0.1

0.33±0.00

0.08±0.002

KM

15.4±0.3

2.2±0.1

0.60±0.01

0.10±0.003

SN

11.3±0.3

2.2±0.2

0.46±0.00

0.14±0.004

WC

17.7±0.3

4.0±0.1

0.50±0.00

0.12±0.002

CB

5.0±0.8

0.9±0.2

0.22±0.02

0.05±0.007

CO

7.2±0.6

0.8±0.1

0.47±0.02

0.09±0.009

CP

3.8±1.4

0.2±0.0

0.24±0.02

0.04±0.011

NB

3.5±0.8

1.6±0.7

0.18±0.02

0.06±0.029

WV

14.4±1.3

1.9±0.4

0.68±0.04

0.06±0.006

Total

10.1±1.1

1.9±0.3

0.43±0.06

0.08±0.009

B. Owner mean estimates (within ecoregion)

Ecoregion

Live Biomass kg C m-2

Dead Biomass kg C m-2

NPP kg C m-2 yr-1

Mortality kg C m-2 yr-1

 

Private

Public

Private

Public

Private

Public

Private

Public

BM

3.7±0.3

6.7±0.1

1.5±0.2

2.0±0.1

0.23±0.01

0.28±0.00

0.04±0.00

0.09±0.01

CR

12.8±0.6

22.2±0.6

3.1±0.2

3.8±0.2

0.81±0.02

0.76±0.01

0.07±0.00

0.07±0.01

EC

5.8±0.4

6.8±0.1

1.5±0.2

1.4±0.1

0.34±0.02

0.33±0.00

0.06±0.01

0.08±0.02

KM

11.1±0.6

15.8±0.3

1.9±0.2

2.2±0.1

0.67±0.02

0.60±0.01

0.07±0.00

0.11±0.01

SN

8.6±0.6

11.6±0.3

1.8±0.2

2.2±0.1

0.46±0.02

0.46±0.01

0.11±0.01

0.14±0.01

WC

9.8±0.8

18.0±0.3

3.7±0.5

4.0±0.1

0.58±0.03

0.50±0.00

0.05±0.00

0.12±0.00

CB

3.7±0.0

5.1±0.8

-

0.9±0.2

0.18±0.00

0.22±0.02

0.05±0.01

0.05±0.01

CO

7.9±0.6

6.4±0.8

1.0±0.2

0.6±0.2

0.54±0.03

0.39±0.03

0.10±0.01

0.06±0.01

CP

3.8±1.0

 

0.2±0.1

-

0.24±0.02

-

0.04±0.00

-

NB

4.0±1.0

3.4±0.8

-

1.6±0.7

0.18±0.04

0.18±0.03

0.02±0.00

0.07±0.01

WV

12.5±1.0

19.0±0.3

1.6±0.5

2.4±0.9

0.67±0.05

0.70±0.06

0.06±0.01

0.06±0.02

Total

7.6±0.2

11.5±0.1

2.1±0.1

2.1±0.0

0.45±0.01

0.44±0.01

0.08±0.01

0.06±0.01

C. Forest type mean estimates (within ecoregion)

Ecoregion

Live Biomass kg C m-2

 

1

2

3

4

5

6

7

All

BM

7.8±0.2

8.5±0.6

1.8±0.2

9.7±0.5

5.8±0.1

2.6±0.0

1.8±0.1

5.4±0.3

CR

21.0±0.7

20.0±1.6

-

26.2±4.0

5.1±1.1

14.3±0.6

2.3±0.8

14.9±1.5

EC

10.4±0.3

5.9±1.6

4.6±0.6

16.7±0.0

5.8±0.1

8.9±1.6

1.6±0.1

7.7±0.8

KM

17.3±0.4

10.4±3.0

-

7.9±0.0

5.4±0.5

14.2±0.5

1.7±0.2

9.5±0.8

SN

14.0±0.2

-

9.3±2.0

-

6.8±0.3

11.5±0.6

1.4±0.1

8.6±0.6

WC

19.0±0.3

20.0±3.0

-

15.0±1.6

7.5±0.4

8.1±1.1

2.1±0.3

12.0±1.1

CB

7.1±3.5

-

-

-

5.8±0.9

-

1.4±0.1

4.8±1.5

CO

8.9±1.3

26.0±0.0

-

-

4.6±0.7

7.1±0.6

-

11.6±0.9

CP

4.0±0.0

-

1.2±0.0

-

6.1±0.0

-

-

3.8±0.0

NB

7.9±4.0

-

2.1±0.4

-

4.2±1.0

-

0.9±0.7

3.8±1.4

WV

16.0±1.5

-

-

-

-

11.0±2.0

5.3±2.4

10.6±2.0

Ecoregion

Dead Biomass kg C m-2

 

1

2

3

4

5

6

7

All

BM

2.3±0.1

3.5±0.4

0.3±0.1

3.4±0.3

1.3±0.1

-

2.1±0.4

2.2±0.2

CR

3.8±0.2

4.5±0.8

-

3.2±0.6

1.4±1.2

2.8±0.2

1.9±0.4

2.9±0.5

EC

2.1±0.1

1.6±0.3

0.68±0.2

4.5±0.0

1.1±0.0

1.6±0.3

1.0±0.1

1.8±0.2

KM

2.4±0.1

1.7±0.1

-

1.4±0.0

1.1±0.1

1.6±0.1

1.0±0.2

1.5±0.2

SN

2.5±0.1

-

1.4±0.5

-

1.3±0.1

1.5±0.2

0.9±0.1

1.5±0.2

WC

4.2±0.1

6.3±1.2

-

4.7±0.9

1.5±0.1

2.9±0.5

3.4±0.8

3.8±0.6

CB

1.5±0.0

-

-

-

0.9±0.2

-

0.2±0.0

0.9±0.2

CO

1.0±0.2

0.8±0.0

-

-

0.7±0.3

0.6±0.1

-

0.8±0.2

CP

-

-

-

-

-

0.2±0.0

-

0.2±0.0

NB

2.7±1.0

-

0.1±0.0

-

0.7±0.5

-

-

1.2±0.8

WV

2.2±0.5

-

-

-

-

0.7±0.2

1.4±0.0

1.5±0.4

Ecoregion

NPP kg C m-2 yr-1

 

1

2

3

4

5

6

7

All

BM

0.29±0.00

0.28±0.014

0.13±0.01

0.28±0.02

0.31±0.00

0.19±0.00

0.12±0.00

0.23±0.01

CR

0.75±0.01

0.92±0.04

-

0.75±0.04

0.41±0.08

0.84±0.02

0.19±0.04

0.65±0.04

EC

0.46±0.01

0.31±0.04

0.21±0.02

0.27±0.00

0.30±0.00

0.56±0.09

0.13±0.00

0.32±0.03

KM

0.60±0.01

0.45±0.07

-

0.26±0.00

0.31±0.02

0.78±0.02

0.14±0.01

0.42±0.02

SN

0.52±0.01

-

0.31±0.05

-

0.34±0.01

0.57±0.02

0.13±0.01

0.37±0.02

WC

0.52±0.01

0.54±0.04

-

0.39±0.04

0.36±0.01

0.48±0.05

0.14±0.01

0.40±0.03

CB

0.26±0.15

-

-

-

0.24±0.02

-

0.11±0.01

0.21±0.06

CO

0.48±0.07

0.87±0.00

-

-

0.33±0.04

0.51±0.04

-

0.54±0.05

CP

0.30±0.00

-

0.10±0.00

-

0.32±0.00

-

-

0.24±0.00

NB

0.36±0.11

-

0.13±0.01

-

0.21±0.03

-

0.08±0.01

0.19±0.04

WV

0.71±0.04

-

-

-

-

0.67±0.10

0.34±0.12

0.57±0.09

Ecoregion

Mortality kg C m-2 yr-1

 

1

2

3

4

5

6

7

All

BM

0.13±0.00

0.12±0.01

0.02±0.00

0.14±0.01

0.06±0.00

0.06±0.00

0.02±0.00

0.08±0.01

CR

0.06±0.00

0.08±0.01

-

0.06±0.01

0.04±0.01

0.08±0.00

0.01±0.00

0.05±0.01

EC

0.16±0.01

0.04±0.01

0.03±0.01

0.16±0.00

0.05±0.00

0.07±0.01

0.01±0.00

0.08±0.01

KM

0.12±0.00

0.07±0.01

-

0.06±0.00

0.04±0.00

0.08±0.00

0.01±0.00

0.06±0.01

SN

0.18±0.01

-

0.07±0.02

-

0.06±0.00

0.11±0.01

0.02±0.00

0.09±0.01

WC

0.13±0.00

0.14±0.02

-

0.15±0.02

0.07±0.01

0.05±0.01

0.01±0.00

0.09±0.01

CB

0.08±0.06

-

-

-

0.05±0.01

-

0.01±0.00

0.05±0.02

CO

0.06±0.01

0.03±0.00

-

-

0.03±0.01

0.11±0.02

-

0.04±

CP

0.04±0.00

-

0.01±0.00

-

0.06±0.00

0.06±0.00

-

0.04±0.00

NB

0.32±0.17

-

0.02±0.00

-

0.03±0.01

0.01±0.00

0.01±0.00

0.08±0.05

WV

0.06±0.01

-

-

-

-

0.06±0.01

0.02±0.01

0.05±0.01

1. Forest Types: 1 =Fir/Douglas fir/Hemlock, 2=Cedar/Larch/Sequoia/Redwood, 3=Juniper/Cypress, 4=Spruce, 5=Pine, 6=Hardwoods, 7=Non-Stocked


TABLE A3. Comparison of ecoregion mean (kg C m-2) and total (Pg C) estimates of live biomass for single condition (SC) and all plots.

Ecoregion

SC Mean

All (mean)

SC (total)

All (total)

Adj. R2*

BM

6.4        

8.6

0.182

0.274

0.75

CR

19.1

18.2

0.627

0.597

0.90

EC

6.8

7.3

0.200

0.218

0.90

KM

15.4

15.4

0.627

0.627

0.90

SN

11.3

12.4

0.347

0.380

0.83

WC

17.7

19.1

0.511

0.551

0.91

CB

5.0

5.9

0.008

0.010

0.99

CO

7.2

7.6

0.128

0.135

0.70

CP

3.8

4.6

0.002

0.002

-

NB

3.5

4.3

0.006

0.008

0.81

WV

14.4

13.3

0.728

0.067

0.88

Total

10.1

10.6

2.71

2.84

 

*Correlation of 5-yr bin means of single condition plots vs. all plots

 

TABLE A4. Methods and equations for computing biomass and NPP components and regional and total stocks and flux.

Component

Code

Equation

Notes

Bole Biomass

Biomassb

Bole Volume Equation*Wood Density1

Bole volume derived from allometric equation using DBH (diameter at breast height) and height

Branch biomass

Biomassbr

Allometric Equation

Derived from DBH and/or height

Bark biomass

Biomassba

Allometric Equation

Derived from DBH and/or height

Foliage biomass

Biomassf

Allometric Equation

Derived (allometry) from DBH and/or height

Coarse root biomass

Biomasscr

Allometric Equation

Derived from a volume equation developed for Douglas-fir and species-specific wood densities

Leaf Area Index

LAI

Biomassf/ LMA

Calculated from Biomassf and leaf mass per unit leaf area (LMA)2

Fine root biomass

Biomassfr

(exp(4.4179+(.3256*LAI)-(.0237*LAI2))

Van Tuyl et al. (2005),  Supplemental plot data (P < 0.001, R2 = 0.41, n = 36)

Understory shrub biomass

Biomasss

a*(1-(exp(-b*Shrub Volume)))4

Shrub volume calculated as the product of the recorded fraction plot cover, plot area, and height

Coarse woody debris

volume

Volumecwd

(9.869/(8*L))*(D2)5

Where, L is the transect length in meters and D is the diameter of the piece in centimeters

Coarse woody debris biomass

Biomasscwd

Volumecwd * Adjusted Density

Derived by multiplying Volumecwd by a decay class adjusted species-specific density

Wood NPP (Bole, Branch, Bark, and Coarse Roots)

NPPw

Biomassw2 – Biomassw1

Difference between biomass of woody components at current and previous time steps

Foliage NPP

NPPf

Biomassf/ Foliage Retention time

Biomass of foliage divided by the average number of years of foliage a stand carries5

Fine root NPP

NPPfr

Biomassfr * 1.2 yearr-1

Average fine root turnover (1.2 year-1) obtained from the literature and supplemental plot data (Keyes and Grier 1981, Campbell et al. 2004a)

Regional estimates

 

Sum of (Ecoregion mean * Forested Area)

 

Theoretical stocks

 

Sum of (Ecoregion asymptote * Forested Area)

Asymptote is parameter ‘a’ from the curve fit to the mean trend of the data (Table 5). 

1 Species-specific wood densities were obtained from US Forest Service wood density survey for western Oregon (Maeglin and Wahlgren 1972), the Forest Products Laboratory wood handbook (1974), and from wood cores obtained on our supplemental plots. Wood densities were reduced according to decay class for standing dead trees (Waddell 2002)

2 LMA was obtained from a look-up table of species-specific values obtained from measurements on the supplemental plots in each of the ecoregions. In some cases, a species-specific value was not available and therefore a closely related species was used.

3 The parameters "a" and "'b" are regression coefficients that vary by species. Equations were developed from harvested shrubs at the supplemental field plots.

4 Equation described in Waddell (2002), Harmon and Sexton (1996) and Van Wagner (1968).

5 Foliage (branch) samples from evergreen species were collected at all supplemental plots and the average number of years of growth retained on each branch was recorded to calculate retention time. Samples were also dried and weighed. New shoot growth was recorded for foliage NPP. This information was used to construct species and ecoregion specific lookup tables for the FIA plots


TABLE A5. Equation1 parameters fit to the mean values of each 5 year age bin.

Live Biomass

Maximum (A)

Rate (B)

Shape (C)

Years to Max2

 

Adj. R2

BM

7.1±0.2

0.028 ±0.008

7.20 ±5.21

180

 

        0.61

CR

33.4±3.0

0.007 ±0.004

0.87 ±0.42

310

 

0.39

EC

10.1±0.8

0.008 ±0.003

1.10 ±0.44

310

 

0.49

KM

44.2±33.0

0.001 ±0.002

0.62 ±0.18

600+

 

0.63

SN

19.4±0.7

0.004 ±0.003

0.81 ±0.31

500

 

0.47

WC

26.9±2.0

0.005 ±0.002

0.86 ±0.20

430

 

0.69

NPP

Maximum (A)

Rate (B)

Decline Age (C)

     

BM

0.30 ±0.01

1.30 ±0.10

216 ±14

125

 

0.57

CR

0.82 ±0.03

2.29 ±0.38

162 ±32

60

 

0.17

EC

0.38 ±0.01

2.39 ±0.46

458 ±197

155

 

0.38

KM

0.65 ±0.01

2.37 ±0.35

324 ±75

110

 

0.31

SN

0.51 ±0.01

1.92 ±0.22

291 ±45

125

 

0.42

WC

0.54 ±0.01

2.09 ±0.15

162 ±12

65

 

0.39

Dead

Maximum

Rate (B)

Shape (C)

Initial

Decay Rate

 

BM

1.39 ±1.10

0.05 ±0.04

276 ±1576

1.83 ±0.9

0.01±0.01

0.15

CR

-

-

-

-

-

-

EC

1.68 ±0.10

0.02 ±0.01

2.51 ±0.01

5.46 ±2

0.05±0.03

0.12

KM

1.57 ±1.93

0.015 ±0.02

33.00 ±166

1.59 ±0.30

0.00 ±0.00

0.24

SN

2.80 ±0.17

0.01 ±0.01

1.61 ±.93

-

-

0.35

WC

9.54 ±5.19

0.004 ±0.004

3.81 ±5.31

3.26 ±1.00

0.003 ±0.01

0.53

Mortality

Maximum (A)

Rate (B)

Shape (C)

Years to Max2

   

BM

0.10 ±0.00

0.035 ±0.015

23.2 ±33.00

140

 

0.48

CR

0.09 ±0.01

0.006 ±0.005

0.48 ±0.27

270

 

0.24

EC

0.11 ±0.01

0.007 ±0.006

1.43 ±1.00

270

 

0.26

KM

0.29 ±0.27

0.001 ±0.003

0.61 ±0.23

600+

 

0.50

SN

0.20 ±0.02

0.008 ±0.004

1.13 ±0.62

250

 

0.34

WC

0.17 ±0.01

0.008 ±0.002

1.20 ±0.34

310

 

0.64

1Equation Forms: Live Biomass = A*(1 - exp(-B*SA)^C, Dead Biomass = D*exp(-E*SA) + A*(1 - exp(-B*SA))^C, NPP = A*exp(-.5*(ln(SA/C)/B)^2), Mortality =  A*(1 - exp(-B*SA)^C where SA is stand age

2Equation solved for the stand age where A is 90% of maximum

TABLE A6. Equation1 parameters fit to upper bounds (99th percentile) of data.

Live Biomass

Maximum (A)

Rate (B)

Shape (C)

Years to Max2

 

Adj. R2

BM

17.8 ±0.9

0.013 ±0.005

1.44 ±0.67

215

 

0.51

CR

70.8 ±15.8

0.004 ±0.004

0.58 ±0.24

445

 

0.44

EC

22.5 ±1.7

0.009 ±0.004

1.01 ±0.46

255

 

0.39

KM

49.1 ±1.9

0.010 ±0.003

1.14 ±0.34

245

 

0.39

SN

36.3 ±1.8

0.014 ±0.006

1.61±0.89

195

 

0.69

WC

53.2 ±1.8

0.008 ±0.002

0.93 ±0.21

275

 

0.62

Dead

Maximum (A)

Rate (B)

Shape (C)

Initial (D)

Decay Rate (E)

 

BM

1.4 ±1.1

0.050 ±0.040

276.0 ±1577

1.83 ±0.87

0.004 ±0.008

0.15

CR

-

-

-

-

-

-

EC

4.9 ±0.3

0.030 ±0.020

5.8 ±11.0

17.10 ±5.99

0.050 ±0.020

0.13

KM

5.0 ±6.0

0.020 ±0.020

17.6 ±71.0

4.58 ±1.22

0.001 ±0.005

0.19

SN

-

-

-

-

-

-

WC

18.6 ±3.0

0.007 ±0.004

3.6 ±5.3

13.48 ±4.26

0.008 ±0.009

0.29

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

   

BM

0.53 ±0.02

1.60 ±0.21

176 ±21

75

 

0.28

CR

1.42 ±0.06

2.12 ±0.29

110 ±17

45

 

0.17

EC

0.74 ±0.02

2.10 ±0.46

278 ±86

105

 

0.32

KM

1.40 ±0.04

1.73 ±0.13

144 ±11

65

 

0.12

SN

1.00 ±0.03

1.86 ±0.17

179 ±16

75

 

0.56

WC

1.05 ±0.02

1.83 ±0.09

104 ±6

45

 

0.34

Mortality

Maximum (A)

Rate (B)

Shape (C)

Years to Max

   

BM

0.27 ±0.01

0.020 ±0.010

8.89 ±9.60

225

 

0.39

CR

0.25 ±0.10

0.001 ±.004

0.02 ±0.10

70

 

0.24

EC

0.34 ±0.02

0.020 ±.010

3.58 ±4.05

175

 

0.26

KM

0.37 ±0.02

0.010 ±.005

1.15 ±0.62

245

 

0.11

SN

0.45 ±0.02

0.030 ±.020

3.36 ±3.53

115

 

0.5

WC

0.37 ±0.01

0.010 ±.004

1.38 ±0.55

260

 

0.32

1Equation Forms: Live Biomass = A*(1 - exp(-B*SA)^C, Dead Biomass = D*exp(-E*SA) + A*(1 - exp(-B*SA))^C, NPP = A*exp(-.5*(ln(SA/C)/B)^2), Mortality =  A*(1 - exp(-B*SA)^C where SA is stand age

2Equation solved for the stand age where A is 90% of maximum

TABLE A7. Regression parameters for equations1 fit to mean values of each 5 year bin (maximum of 200 years) by ownership.

Public

Live

Maximum (A)

Rate (B)

Shape (C)

Years to Max2

Adj. R2

BM

11.2±4.4

0.006±0.005

1.04±0.40

390

0.76

CR

27.0±2.7

0.020±0.011

1.75±0.98

145

0.66

EC3

8.8±1.5

0.011±0.009

1.36±0.77

235

0.86

KM

18.1±2.7

0.014±0.006

1.76±0.67

205

0.84

SN

14.0±1.2

0.020±0.006

2.65±1.02

165

0.86

WC

21.2±2.9

0.011±0.005

1.38±0.36

280

0.91

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

 

BM

0.33±0.12

3.16±2.08

250+

250+

0.33

CR

0.81±0.03

1.69±0.23

87±11

45

0.45

EC

0.34±0.02

1.91±0.41

233±115

95

0.70

KM

0.63±0.03

1.50±0.29

165±51

85

0.62

SN

0.58±0.11

2.22±0.63

250+

185

0.78

WC

0.55±0.01

1.31±0.10

96±6

55

0.79

Mortality

Maximum (A)

Rate (B)

Shape (C)

Years to Max

 

BM

0.172±0.076

0.009±0.007

2.14±1.03

335

0.84

CR

0.054±0.002

0.103±0.042

7.41±7.71

45

0.59

EC

0.093±0.013

0.017±0.008

2.07±1.01

105

0.78

KM

0.112±0.014

0.020±0.009

2.48±1.38

155

0.74

SN

0.160±0.010

0.029±0.009

4.05±2.25

125

0.80

WC

0.170±0.040

0.009±0.005

1.23±0.36

280

0.89

Private

Live

Maximum (A)

Rate (B)

Shape (C)

Years to Max

Adj. R2

BM

4.8±0.4

0.05±0.03

31.1±64.8

115

0.57

CR

17.1±1.0

0.04±0.02

3.0±2.2

85

0.67

EC

7.9±1.1

0.03±0.02

6.4±11.0

135

0.32

KM

14.6±2.1

0.02±0.02

2.0±2.1

160

0.36

SN

10.2±1.2

0.03±0.02

4.4±5.8

125

0.55

WC

16.8±6.6

0.01±0.02

1.0±1.2

240

0.42

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

 

BM

0.27±0.02

0.83±0.17

143±24

105

0.60

EC

0.43±0.08

1.30±0.54

245±187

135

0.53

SN

0.52±0.03

0.91±0.20

119±17

75

0.51

CR

0.96±.04

1.45±0.25

121±25

65

0.65

WC

0.66±0.05

1.42±0.44

104±36

55

0.27

KM

0.76±.04

1.63±0.50

100±27

45

0.13

Mortality

Maximum (A)

Rate (B)

Shape (C)

Years to Max

 

BM

0.051±0.005

0.063±0.036

75±221

105

0.63

CR

0.093±0.005

0.155±0.111

152±583

45

0.51

EC

0.081±0.016

0.065±0.064

132±664

175

0.25

KM

0.082±0.010

0.070±0.121

3.6±13.1

55

0.17

SN

0.150±0.031

0.018±0.021

2.5±4.5

175

0.43

WC

0.180±0.081

0.006±0.008

1.6±1.3

250+

0.32

1Equation Forms: Live Biomass = A*(1 - exp(-B*SA)^C, NPP = A*exp(-.5*(ln(SA/C)/B)^2), Mortality =  A*(1 - exp(-B*SA)^C where SA is stand age

2Equation solved for the stand age where A is 90% of maximum

3Data included for age groups up to 250 (live biomass only) for EC and BM because values were still significantly increasing in both ownerships


TABLE A8. Regression parameters for equations1 fit to mean values of each 5 year bin by forest type.  Regressions could not be fit to some forest types in some ecoregions as there were not enough age classes with data.

1: Douglas-fir/Hemock

Live

Maximum (A)

Rate (B)

Shape (C)

Years to Max2

Adj. R2

BM

9.1±0.5

0.023±0.010

5.95±6.55

175

0.31

CR

33.4±2.1

0.012±0.006

1.23±0.66

210

0.41

EC

14.9±2.3

0.006±0.005

0.91±0.52

345

0.34

KM

42.4±24.9

0.001±0.002

0.65±0.21

600+

0.58

SN

17.7±0.9

0.014±0.006

1.77±1.03

205

0.37

WC

28.0±2.0

0.005±0.002

0.78±0.18

420

0.69

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

 

BM

0.32±0.01

1.20±0.19

224±21

130

0.20

CR

0.78±0.03

2.55±0.48

159±34

50

0.12

EC

0.51±0.03

2.25±0.72

472±322

170

0.23

KM

0.64±0.01

2.27±0.34

320±70

120

0.28

SN

0.56±0.01

1.71±0.18

228±24

110

0.31

WC

0.56±0.01

2.03±0.13

149±10

60

0.43

Mortality

Maximum (A)

Rate (B)

Shape (C)

Years to Max

 

BM

0.16±0.01

0.023±0.011

7.13±8.47

185

0.30

CR

0.10±0.02

0.004±0.005

0.53±0.30

430

0.26

EC

0.21±0.02

0.012±0.008

2.05±1.84

250

0.27

KM

0.21±0.05

0.003±0.003

0.74±0.27

600+

0.49

SN

0.21±0.01

0.02±0.010

1.97±1.58

150

0.25

WC

0.17±0.01

0.008±0.002

1.15±0.34

310

0.62

2: Cedar/Larch

Live

Maximum (A)

Rate (B)

Shape (C)

Years to Max

Adj. R2

BM

8.4±0.5

0.044±0.072

2.73±8.00

600+

0.06

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

 

CR

0.97±0.06

1.89±0.47

118±29

50

0.23

Mortality

Maximum (A)

Rate (B)

Shape (C)

Years to Max

 

CR

0.08±0.01

0.169±0.188

249±1605

50

0.22

KM

0.21±0.05

0.003±0.003

0.74±0.27

600+

0.49

3: Juniper

Live

Maximum (A)

Rate (B)2

Shape (C)

Years to Max

Adj. R2

BM

3.0±0.3

0.018±0.009

2.68±2.17

190

0.50

EC

4.5±2.6

0.005±0.050

0.15±0.57

140

0.00

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

 

BM

0.16±0.01

1.56±0.30

233±48

115

0.35

Mortality

Maximum (A)

Rate (B)2

Shape (C)

Years to Max

 

BM

0.03±0.00

0.023±0.018

4.12±5.88

160

0.29

4: Spruce – NONE

5: Pine

Live

Maximum (A)

Rate (B)2

Shape (C)

Years to Max

Adj. R2

BM

9.7±3.9

0.003±0.004

0.54±0.26

580

0.43

EC

9.2±0.9

0.005±0.002

0.75±0.21

410

0.64

KM

8.0±3.7

0.005±0.010

0.61±0.61

370

0.11

SN

8.3±0.4

0.026±0.014

1.84±1.47

120

0.29

WC

8.9±0.4

0.082±0.044

21.91±41.0

70

0.38

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

 

BM

0.32±0.01

2.74±0.97

422±310

120

0.18

EC

0.33±0.01

2.99±0.60

527±283

130

0.43

KM

0.29±0.03

3.41±3.74

170±236

35

0.00

SN

0.35±0.01

2.73±0.80

227±102

65

0.09

WC

0.41±0.02

1.85±0.48

133±32

55

0.07

Mortality

Maximum (A)

Rate (B)2

Shape (C)

Years to Max

 

BM

0.21±1.17

0.001±0.006

0.50±0.32

600+

0.34

EC

0.07±0.00

0.011±0.006

0.95±0.51

240

0.33

KM

0.05±0.01

0.008±0.012

0.72±0.77

260

0.10

SN

0.08±0.01

0.023±0.016

1.76±1.71

130

0.20

WC

0.09±0.01

0.030±0.017

3.26±3.36

120

0.35

6: Hardwoods

Live

Maximum (A)

Rate (B)2

Shape (C)

Years to Max

Adj. R2

CR

30.0±6.3

0.004±0.003

0.67±0.23

500

0.58

KM

23.1±2.1

0.007±0.004

1.06±0.43

300

0.51

SN

13.8±0.8

0.035±0.014

9.71±10.71

130

0.50

WC

9.7±1.2

0.137±0.208

95±677

50

0.18

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

 

CR

0.98±0.67

2.48±0.78

355±265

115

0.28

EC

0.70±0.12

1.08±0.49

231±85

145

0.10

KM

0.90±0.03

1.94±0.38

294±87

125

0.27

SN

0.65±0.02

1.44±0.24

280±59

145

0.52

WC

0.72±0.07

1.00±0.16

90±13

55

0.45

Mortality

Maximum (A)

Rate (B)

Shape (C)

Years to Max

 

KM

0.11±0.01

0.010±0.007

0.92±0.57

230

0.26

SN

0.13±0.01

0.035±.018

6.51±8.10

120

0.35

7: Non-Stocked

Live

Maximum (A)

Rate (B)

Shape (C)

Years to Max

Adj. R2

BM

2.8±0.4

0.007±0.004

1.16±0.51

350

0.65

EC

2.3±0.3

0.010±0.007

0.90±0.53

230

0.36

KM

3.4±2.8

0.004±0.015

0.67±0.89

490

0.24

SN

2.0±0.3

0.019±0.019

1.20±1.38

140

0.25

NPP

Maximum (A)

Rate (B)

Decline Age (C)

Years to Max

 

BM

0.13±0.01

1.59±0.32

231±44

125

0.27

EC

0.15±0.01

2.13±0.58

180±60

70

0.08

SN

0.14±0.01

1.82±0.38

128±29

55

0.22

Mortality

Maximum (A)

Rate (B)2

Shape (C)

Years to Max

 

BM

0.37±0.01

0.007±0.006

1.34±0.887

370

0.46

EC

0.03±0.04

0.002±0.009

0.474±0.477

600+

0.13

SN

.03±.01

0.058±0.050

19.07±53.55

230

0.24

1Equation Forms: Live Biomass = A*(1 - exp(-B*SA)^C, NPP = A*exp(-.5*(ln(SA/C)/B)^2), Mortality =  A*(1 - exp(-B*SA)^C where SA is stand age

2Equation solved for the stand age where A is 90% of maximum


TABLE A9. Top ranked explanatory variables (from model weights) across entire study region (ecoregion, ownership, or forest type) and within each ecoregion (ownership or forest type), after accounting for age.

Region

Live Biomass

Dead Biomass

NPP

Mortality*

Study Area

Forest Type

Ecoregion

Ecoregion

Forest Type

BM

Forest Type

Forest Type

Forest Type

Forest Type

CR

Forest Type

Forest Type

Forest Type

Owner

EC

Forest Type

Forest Type

Forest Type

Forest Type

KM

Forest Type

Forest Type

Forest Type

Forest Type

SN

Forest Type

Forest Type

Forest Type

Forest Type

WC

Forest Type

Forest Type

Forest Type

Forest Type

*Effects of each factor on mortality could only be assessed for stands <= 250 years because transformations failed to adjust for non-normal distribution of all data points.

LITERATURE CITED

Campbell, J. L., O. J. Sun, and B. E. Law. 2004. Supply-side controls on soil respiration among Oregon forests. Global Change Biology 10:1857–1869.

Forest Products Laboratory. 1974. Wood Handbook: Wood as an engineering material. Forest Products Laboratory, Madison, WI, p. 72.

Harmon, M. E., and J. Sexton. 1996. Guidelines for Measurements of Woody Debris in Forest Ecosystems. Publication No. 20. US LTER Network Office, University of Washington, Seattle, WA:93pp.

Keyes, M. R., and C. C. Grier. 1981. Above- and belowground net production in 40-year-old Douglas-fir stands on low and high productivity sites. Canadian Journal of Forest Research 11:599–605.

Maeglin, R. R., and H. E. Wahlgren. 1972. Western Wood Density Survey: Report Number 2. USDA Forest Service Research Paper FPL-183.

Van Tuyl, S., B. E. Law, D. P. Turner, and A. I. Gitelman. 2005. Variability in net primary production and carbon storage in biomass across Oregon forests--an assessment integrating data from forest inventories, intensive sites, and remote sensing. Forest Ecology and Management 209:273–291.

Van Wagner, C. E. 1968. The line-intersect method in furest fuel sampling. Forest Science 14:20–26.

Waddell, K. L. 2002. Sampling coarse woody debris for multiple attributes in extensive resource inventories. Ecological Indicators 1:139–153.


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