Ecological Archives A022-006-A1

Kyla M. Dahlin, Gregory P. Asner, and Christopher B. Field. 2012. Environmental filtering and land-use history drive patterns in biomass accumulation in a mediterranean-type landscape. Ecological Applications 22:104–118.

Appendix A (TABLE A1). A list of species-specific and generic allometric equations used to calculate aboveground biomass.

Allometric Equation




Sources Species/ Type


JRBP Species Applied to

Equation Source Document

Acacia auriculiformis

BAT = 3.82 + 1.56*DBH

Acacia spp.

Lieurance, 2007

Acer negundo

BAT = .117+.01659*(DBH^2*HT)+0.161+.0007573*(DBH^2*HT)

Acer negundo v. californicum

Schlaegel, 1982

Adenostoma fasciulatum

BAT = 0.18*(pi()(DBA/2)^2) + 0.06

Adenostoma fasciulatum

Stoehlgren, 1982

Arbutus menziesii

BAT = (.0000821921*(DBH^1.96628)*(HT^0.83458))*572

Arbutus menziesii

Pillsbury, 1984 and Cornwell, 2006

Arctostaphylos patula

BAT = (exp( 3.3186 + 2.6846*ln(DBA)))/1000

Arctostaphylos tomentosa*

Hopkins and Busse, 1992#

Artemisia tridentata

BAT = (exp(3.161 + 2.242 * ln(DBA)))/1000

Artemisia californica*

Brown, 1976

Baccharis pilularis*

dead - unknown (shrubs)

Eriodictyon californicum

Genista monspessulana

Holodiscus discolor*

Lepechinia calycina


BAT = exp(-2.2094+2.3867*ln(DBH))

Alnus rhombifolia

Jenkins et al, 2003

Ceanothus velutinus

BAT = (exp(3.670 + 2.847 * ln(DBA)))/1000

Ceanothus cuneatus var cun.*

Brown, 1976

Ceanothus oliganthus*

Heteromeles arbutifolia*

Prunus ilicifolia ssp. Ilicifolia (shrubs)

Quercus durata var. durata

Rhamnus californica*

Rhamnus crocea*

Rhamnus tomentella*

Cedar / Larch

BAT = exp(-2.0336+2.2592*ln(DBH))

Sequoia sempervirens

Jenkins et al, 2003

Cornus sericea%

BAT = (exp(3.681 + 2.575 * ln(DBA)))/1000

Cornus sericea

Brown, 1976

Dirca occidentalis

Corylus cornuta var. ca

BAT = (0.33396 * DBA^2 * HT +  exp(2.4170 + 2.040  * ln(DBA)))/1000

Corylus cornuta var. ca

Buckman, 1966# and Gholz, 1979#

Lonicera utahensis

BAT = (exp(3.768 + 2.957 * ln(DBA)))/1000

Clematis lasiantha

Brown, 1976

Lonicera hispidula var. vac.*

Lonicera interrupta*

Sambucus mexicana* (shrubs)

Toxicodendron diversilobum*

Mimulus guttatus

BAT = 0.981 * DBA * .001

Mimulus aurantiacus

Koerper, 1983

Mixed hardwoods

BAT = exp(-2.48+2.4835*ln(DBH))

Aesculus californica

Jenkins et al, 2002

dead - unknown (trees)

Juglans californica var. hindsii

Prunus ilicifolia ssp. Ilicifolia (trees)


BAT = exp(-2.5356+2.4349*ln(DBH))

Pinus radiata

Jenkins et al, 2003

Quercus agrifolia

BAT = (.0000446992*(DBH^2.31958)*(HT^0.62528))*700

Quercus agrifolia

Pillsbury, 1984 and Cornwell, 2006

Quercus douglasii

BAT = (.0000697541*(DBH^2.33089)*(HT^0.461))*755

Quercus douglasii

Pillsbury, 1984 and Cornwell, 2006

Quercus kelloggii

BAT = (.0000870843*(DBH^1.97437)*(HT^0.85034))*722

Quercus kelloggii

Pillsbury, 1984 and Cornwell, 2006

Quercus lobata

BAT = (.0000334750*(DBH^2.33631)*(HT^0.74872))*705

Quercus lobata

Pillsbury, 1984 and Cornwell, 2006

Ribes spp.

BAT = (exp(2.805 + 2.504 * ln(DBA)))/1000

Ribes spp.

Halpern and Miller, 1993

Rosa spp.

BAT = (exp(3.628 + 2.779 * ln(DBA)))/1000

Rosa spp.

Brown, 1976

Salix spp.

BAT = (exp (3.303+2.762*ln(DBA)))/1000

Salix lasiolepis

Brown, 1976

Salix lucida ssp. lasiandra

Salix sitchensis

Symphoricarpos albus

BAT = (exp(3.490 + 2.285 * ln(DBA)))/1000

Symphoricarpos albus var. laevigatus

Brown, 1976

Umbellularia californica

BAT = (.0000763133*(DBH^1.94553)*(HT^0.88389))*515

Umbellularia californica

Pillsbury, 1984 and Cornwell, 2006


BAT = exp(-0.7152+1.7029*ln(DBH))

Cercocarpus betuloides v. bet**

Jenkins et al, 2003

Olea europea



Sambucus mexicana (trees)



BAT = total aboveground biomass; DBH = diameter at breast height; DBA = stem basal diameter; HT = tree height

In cases where dbh was needed but dba was measured, dba was multiplied by 0.9

In cases where dba was needed but dbh was measured, dbh was multiplied by 1.1

In the ONE case where percent cover was needed (Mimulus), DBA was multiplied by .001 to simulate percent cover.

In equations where height was required, heights were estimated using lidar - trees noted as "upper" canopy were assigned the maximum height measured in the plot, trees noted as "mid" canopy were assigned max ht * 0.5, trees noted as "lower" were all designated as 2.5 m tall. Shrubs were designated as 1.5 m tall.

Bold "Source Species / Types" are categories from Jenkins et al., 2003, applied only where no other appropriate equation could be found.

Original Pillsbury (1984) equations were for volume in cubic meters, so they are converted to biomass by multiplying by wood density from Preston et al (2006) in kg m-3

* = Species equivalence (or or a congener) used in Riccardi et al. 2007

** = Equation applied to congeners in Jenkins 2002

% = Listed in BIOPAK as C. stolonifera, but these species are the same and C. sericea is current according to

# = source from BIOPAK (Means et al. 1994)


Brown, J. K. 1976. Estimating shrub biomass from stem basal diameters. Canadian Journal of Forest Research 6:153–158.

Buckman, R. E. 1966. Estimation of cubic volume of shrubs (Corylus spp.) Ecology 47(5): 858–860.

Gholz, H. L., C. C. Grier, A. C. Campbell, and A. T. Brown.1979. Equations for estimating biomass and leaf area of plants in the Pacific Northwest.  Oregon State University, Forest Research Laboratory, Research Paper 41. 39 pp.

Hopkins, W., and M. Busse. 1992. Unpublished data from BIOPAK (Means et al 1994).

Jenkins, J. C., D. C. Chojnacky, L. S. Heath, and R. A. Birdsey. 2003. National-scale biomass estimators for United States tree species. Forest Science. 49(1):12–35.

Koerper, G. 1983. Unpublished data from BIOPAK (Means et al 1994).

Lieurance, D. M. 2007. Biomass allocation of the invasive tree Acacia auriculiformis and refoliation following hurricane-force winds. Journal of the Torrey Botanical Society 134(3):389–397.

Means, J. E., H. A. Hansen, G. J. Koerper,P. B. Alaback, and M. W. Klopsch. 1994. Software for computing plant biomass--BIOPAK users guide. USDA Forest Service, Pacific Northwest Research Station, General Technical Report 340.

Preston, K. A., W. K. Cornwell, and J. L. DeNoyer. 2006. Wood density and vessel traits as distinct correlates of ecological strategy in 51 California coast range angiosperms. New Phytologist 170:807–818.

Schlaegel, B. E. 1982. Boxelder (Acer negundo L.) biomass component regression analysis for the Mississippi Delta. Forest Science 28(2):355–358.

Stoehlegren, T. J., N. L. Stephenson, D. J. Parsons, and P.W. Rundel. 1982. Using stem basal area to determine biomass and stand structure in chamise chaparral. Presentation at the Symposium on Dynamics and Management of Mediterranean-type Ecosystems, June 22-26, San Diego, California, USA.

Pillsbury, N. H. 1984. Equations for total, wood, and saw-log volume for thirteen Californa hardwoods. USDA Forest Service, Pacific Northwest Research Station, Reasearch Note 414.

[Back to A022-006]