Ecological Archives E090-071-A1

Mark C. Drever, Jacob R. Goheen, and Kathy Martin. 2009. Species–energy theory, pulsed resources, and regulation of avian richness during a mountain pine beetle outbreak. Ecology 90:1095–1105.

Appendix A (Table A1). A list of species seen at 10 forest sites in interior British Columbia, 1997–2007. Species are listed by foraging guild, and an asterisk indicates that the species is known to forage on adult and/or larval forms of the mountain pine beetle (Dendroctonus ponderosae) (from Martin et al. 2006).

Aerial insectivore

Common nighthawk (Chordeiles minor)

Olive-sided flycatcher (Contopus cooperi)*

Western wood-pewee (Contopus sordidulus)*

Alder flycatcher (Empidonax alnorum)

Dusky/Hammond's flycatcher (Empidonax oberholseri/hammondii)

Least flycatcher (Empidonax minimus)

Pacific-slope flycatcher (Empidonax difficilis)*

Willow flycatcher (Empidonax traillii)

Western kingbird (Tyrannus verticallis)

Red-eyed vireo (Vireo olivaceus)

Barn Swallow (Hirundo rustica)

Northern rough-winged swallow (Stelgidopteryx serripennis)

Tree swallow (Tachycineta bicolor)*

Violet-green swallow (Tachycineta thalassina)*

Townsend's solitaire (Myadestes townsendi)

Bark insectivore

Red-naped sapsucker (Sphyrapicus nuchalis)*

American Three-toed woodpecker (Picoides dorsalis)*

Black-backed woodpecker (Picoides arcticus)*

Downy woodpecker (Picoides pubescens)*

Hairy woodpecker (Picoides villosus)*

Pileated woodpecker (Dryocopus pileatus)*

Red-breasted nuthatch (Sitta canadensis)*

Brown creeper (Certhia americana)*

Foliage insectivore

Cassin's Vireo (Vireo cassinii)

Warbling vireo (Vireo gilvus)

Black-capped chickadee (Poecile atricapillus)*

Boreal chickadee (Poecile hudsonica)*

Mountain chickadee (Poecile gambeli)*

Golden-crowned kinglet (Regulus satrapa)

Ruby-crowned kinglet (Regulus calendula)

Common yellowthroat (Geothlypis trichas)

MacGillivray's warbler (Oporornis tolmiei)

Orange-crowned warbler (Vermivora celata)

Townsend's warbler (Dendroica townsendi)

Wilson's warbler (Wilsonia pusilla)

Magnolia warbler (Dendroica magnolia)

Yellow-rumped warbler (Dendroica coronata)*

Yellow warbler (Dendroica petechia)

Blackpoll warbler (Dendroica striata)

Western tanager (Piranga ludoviciana)*

Ground insectivore

Common snipe (Gallinago gallinago)

Northern flicker (Colaptes auratus)*

Marsh wren (Cistothorus palustris)

Winter wren (Troglodytes troglodytes)

Mountain bluebird (Sialia currucoides)*

Hermit thrush (Catharus guttatus)

Swainson's thrush (Catharus ustulatus)

Veery (Catharus fuscescens)

American robin (Turdus migratorius)*

Varied thrush (Ixoreus naevius)

European starling (Sturnus vulgaris)

American pipit (Anthus rubescens)

Northern waterthrush (Seiurus noveboracensis)

Clay-colored sparrow (Spizella pallida)

Chipping sparrow (Spizella passerina)

Vesper sparrow (Pooecetes gramineus)

Savannah sparrow (Passerculus sandwichensis)

Lincoln's sparrow (Melospiza lincolnii)

Song sparrow (Melospiza melodia)

Golden-crowned sparrow (Zonotrichia atricapilla)

White-crowned sparrow (Zonotrichia leucophrys)

Red-winged blackbird (Agelaius phoeniceus)

Western meadowlark (Sturnella neglecta)

Yellow-headed blackbird (Xanthocephalus xanthocephalus)

Brewer's blackbird (Euphagus cyanocephalus)

Rusty blackbird (Euphagus carolinus)

Brown-headed cowbird (Molothrus ater)


Ruffed grouse (Bonasa umbellus)

Sharp-tailed grouse (Tympanuchus phasianellus)

Rufous hummingbird (Selasphorus rufus)

Evening grosbeak (Coccothraustes vespertinus)

Bohemian waxwing (Bombycilla garrulus)

Cedar waxwing (Bombycilla cedrorum)

Dark-eyed junco (Junco hyemalis)*

Pine grosbeak (Pinicola enucleator)

Cassin's finch (Carpodacus cassinii)

Purple finch (Carpodacus purpureus)

Red crossbill (Loxia curvirostra)

White-winged crossbill (Loxia leucoptera)

Pine siskin (Carduelis pinus)


Gray jay (Perisoreus canadensis)

Black-billed magpie (Pica hudsonia)

American crow (Corvus brachyrhynchos)

Common raven (Corvus corax)


Martin, K., A. Norris, and M. Drever. 2006. Effects of bark beetle outbreaks on avian biodiversity in the British Columbia interior: Implications for critical habitat management. BC Journal of Ecosystems and Management 7:10–24. URL:

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